Epaulet Ladybird Rhyzobius chrysomeloides
Other names: Arboreal Ladybird Round-keeled Rhyzobius
This is the commonest ladybird in my recording area, occurring in pine trees, Ivy and ornamental evergreen shrubs.
It is also the only species that I can find, almost at will, on any day of the year.
It occurs in every local garden with ornamental shrubs that I have searched.
It can easily be found wintering in Ivy, using a beating tray. A useful project during the quiet months of November-February.
The colour contrast between the light and dark markings show up best on photographs of live specimens taken outside in natural light, using a digital camera with a macro-lens. Studio photographs of pinned specimens tend to bleach out much of the subtle colour details.
Identification Length 2.5-3.5mm
A brown, long-bodied oval species with long antennae.
Epaulet Ladybird has a reputation for being easily confused with Meadow Ladybird but I consider the majority to be identifiable in the field.
The ground colour varies from light oak to deep chestnut.
An obvious feature is a pale curved stripe across the top of the wing cases.
This is sometimes reduced to an orange patch on the shoulder areas, resembling epaulets.
In the most distinctive individuals the curved stripe continues as two stripes running either side of the wing case centre line, forming a pair of tramlines.
The outer wing cases typically have a shield shaped area of dark lines surrounding the pale shoulder curve and tramlines.
This usually includes two vertical lines on each wing case, although these can be fused or broken.
The outer edge has a distinctive curved S-shape, with a scalloped indentation at the top and a bulge towards the rear.
The centre rear often has an anchor shaped mark, with a tooth-like projection on each side contrasting with a pale area on the wing case tips.
The pronotum (between the head and wing cases) have straight and nearly parallel sided edges to the rear.
Wing case pattern
The most distinctive individuals have a pale curved stripe running from shoulder to shoulder with a pair of pale tramlines running down either side of the wing case centre.
This common pattern is very obvious and eye catching and is often the form that attracts an observer when first discovering this species. These are unlike any Meadow Ladybirds and are very easy to identify.
Very distinctive pale central lines diverging slightly
Upshire, North Essex
Central lines running parallel to each other
Upshire, North Essex
Waltham Abbey, North Essex
Turnford, Lee Valley, Hertfordshire
Waltham Abbey, North Essex
On some the pale tramlines are broken or irregular.
Tramlines separated from shoulder stripe by dark markings
Sawbridgeworth, Hertfordshire
Indistinct broken pattern Broxbourne, Hertfordshire
Some have a brighter orange patch, resembling an epaulet, on the outer side of the curved shoulder stripe.
Waltham Abbey, North Essex
Fishers Green, North Essex
A small number have the orange patches merging and extending over the top part of the wing cases.
Upshire, North Essex
Sometimes the two tramlines are fused into an extra wide central pale area continuing downwards from the top orange band.
Upshire, North Essex
Some Epaulet Ladybirds are mostly dark but unlike the darkest Meadow Ladybirds they are not uniformly dark.
There is usually a ghostly hint of the pale shoulder curve and the pair of tramlines running down either side of the wing case centre line.
The typical dark markings are also usually present, although often obscured.
The dark tooth-like projections often contrast with the pale, often grey, wing case tips.
Dark with pale shoulder patches
Waltham Abbey, North Essex
Dark with hint of epaulet and central lines pattern
Fishers Green, Lee Valley, North Essex
Overlying the light markings is a distinctive, though variable, dark area that is broadly shield shaped.
The typical pattern has two vertical lines on each wing case.
There is often an anchor shaped mark at the centre rear with a dark line running up the centre.
The outer edge has a scalloped shape near the top, with a bulge further down ending in a tooth shaped point contrasting with the pale rear .
Upshire, North Essex
Typical shield shaped dark markings with double outer line
Exeter, South Devon
Anchor shape extended and resembling Norman helmet with nose guard Exeter, South Devon
On some the black markings are fused together creating a solidly dark shield shaped area on the rear wing cases.
Upshire, North Essex
Upshire, North Essex
The trickiest Epaulet Ladybirds to identify have the dark markings reduced.
However there are always a few clues. The broken dark markings still tend to cover the usual template, with the distinctive outer S-shape (with upper scallop and lower bulge) and usually at least a hint of the second inner line.
Meadow Ladybird only has a single vertical line on each wing case, which is straighter and usually stops at the half way point.
Upshire, North Essex
Upshire, North Essex
Upshire, North Essex
Fishers Green, Lee Valley, North Essex
Very reduced markings but second inner line still present
Waltham Abbey, North Essex
A small number lack the second inner line and these can be very similar to some Meadow Ladybirds, although there are still a few useful features that often help.
The outside shape of the black line is often distinctive with the S-shape formed by the scalloped upper part and bulge on the lower part, with the whole line stretching towards the top corner. I think the presence of the black line in the top corner is likely to be an important identification feature and might prove to be a diagnostic, I have never seen this on Meadow Ladybird.
The central panel is often a bright orange colour, with a series of thin darker horizontal lines.
On darker brown individuals there is often a hint of the brighter orange shoulder patch or epaulet.
The typical straight sided pronotum edge is also often present.
Upshire, North Essex
Upshire, North Essex
Upshire, North Essex
Upshire, North Essex
Waltham Abbey, North Essex
Upshire, North Essex
Recently emerged individuals are a pale fawn or orange colour with faint markings that when looked at carefully still follow the typical pattern.
Probably recently emerged with pale markings Waltham Abbey, North Essex
Recently emerged with very pale hint of typical pattern
Upshire, North Essex
When photographing a suspected Epaulet Ladybird, it is best to get a photograph from above, as this shows the pattern most clearly.
A side on photograph will not show a subtle pattern well, although it will still be useful, as
other features may be apparent.
Indistinct dark and light markings
Braunton Burrows, North Devon
The same individual as above, showing hint of the epaulet stripes and tramlines, as well as pale grey wing case tips
Pronotum Shape
The pronotum shape can help in the identification process but is not always diagnostic.
The majority show a strong curve at the corner forming a right angle with a straight edged
rear section.
The rear section is often nearly parallel sided.
Some are more curved and similar to Meadow Ladybird but most show at least a short straight sided rear section.
Meadow Ladybird usually has a gently curved pronotum, lacking the right angled corner and still curving as it reaches the wing cases.
Very straight sided pronotum
Typical pronotum shape
Antennae Shape
The clubbed antennae are thin, longer than the head width and often held towards the side.
They usually show a distinctive kink at the half way point, resembling a wire coat hanger.
This is the same as Meadow Ladybirds and is typical of ladybirds, so is a good way to rule out look alike beetle species from other family groups.
When disturbed Epaulet Ladybird often sit still with their legs and antennae tucked under the body, so need to be left until they start walking about before this feature can be checked.
Abbey Gardens, Waltham Abbey, North Essex
Mixed Photographs
Fishers Green, Lee Valley, North Essex
Upshire, North Essex
Waltham Abbey, North Essex
With 7-spot Ladybird
With 22-spot and 7-spot Ladybirds
With Cream-spot Ladybird
With Cream-streaked Ladybird
With Larch Ladybird
With 10-spot Ladybirds
With 22-spot Ladybird
With Ivy Ladybird
With 22-spot and Forestier's Ladybirds
Beaten from Ivy alongside Opilo mollis and Pogonocherus hispidus Fishers Green, Lee Valley, North Essex
With two Lesser Thorn-tipped Longhorns
On lid of garden wheelie bin Upshire, North Essex
Variation and Forms
Habitat
Epaulet Ladybird can be found in a wide range of habitats.
They are best searched for using a beating tray (or umbrella).
I have found them by eye on and in garden wheelie bins, containing shrub prunings.
Unlike the slightly scarcer Red-flanked, Ivy and Meadow Ladybirds, I have never found this species by eye on vegetation. It is cryptically coloured and may keep out of sight underneath the vegetation.
Epaulet Ladybird can be found through out the year and is often easy to locate during the winter months, except during frosty, windy or rainy weather.
There is sometimes a drop in numbers in the early spring. April can be the trickiest month to find them, presumably the wintering adults have died off and the next generation has yet to emerge.
Numbers then increase during the summer, with this species becoming abundant in the autumn. July to September can be a very good months to search for Epaulet Ladybirds as they spread into a myriad of available habitats.
Epaulet Ladybirds can be common in pine trees, in small copses or individual trees.
Plantation pine tree Upshire, North Essex
Garden pine tree Upshire, North Essex
Small pine tree Braunton Burrows, North Devon
Pine tree May Day Farm, The Brecks, West Suffolk
Pine tree Turnford, Lee Valley, Hertfordshire
Pine copse Harlow, North Essex
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Amongst pine needles Upshire, North Essex
They can also be common on Ivy in woodland situations.
In the Lee Valley and around the northern edge of Epping Forest they can easily be found in winter by searching Ivy growing on tree trunks.
Sheltered green lane with plenty of Ivy; ideal habitat
Puck Lane, Waltham Abbey, North Essex
Ivy. Fishers Green, Lee Valley, North Essex
Ivy Darland Banks, Gillingham, East Kent
Ivy Turnford, Lee Valley, Hertfordshire
Ivy covered tree trunk Leigh Delamere services, North Wiltshire
Ivy Roding Valley Meadows, South Essex
Ivy Roding Valley Meadows, South Essex
Ivy covered building Hooks Marsh, Lee Valley, North Essex
Ivy covered wall Harlow, North Essex
In December 2020 I found Epaulet Ladybirds in the Holly understorey in Epping Forest for the first time and then at Hainault Forest CP in 2021.
I have previously found Orange and Forestier's Ladybirds in this habitat.
Holly Fairmead, Epping Forest, South Essex
Holly Fairmead, Epping Forest, South Essex
Holly Hainault Forest CP, South Essex
Yew is another evergreen shrub that can be successfully searched for Epaulet Ladybirds.
I also occasionally find them in Holm Oak.
Yew Rushy Mead, Lee Valley, North Essex
Yew Hainault Forest CP. South Essex
Yew Ospringe, East Kent
Holm Oak Upshire, North Essex
Epaulet Ladybirds can be abundant in gardens, both rural and urban.
They can be found on Ivy covered fences and on a variety of evergreen shrubs, especially Euonymus, Viburnum tinus, Firethorn, Privet and Cotoneaster.
I have also occasionally found them on Oleaster Elaeagnus and Berberis.
Euonymus. Upshire, North Essex
Euonymus. Waltham Abbey, North Essex
Euonymus hedge. Abbey Gardens, Lee Valley, North Essex
Firethorn. Upshire, North Essex
Oleaster Elaeagnus Waltham Abbey, North Essex
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Privet hedge Exeter, South Devon
Epaulet Ladybirds can easily be found in garden wheelie bins that have been filled with the prunings from these shrubs.
They can also be found in deciduous shrubs, including Weigela, Dogwood and Guelder-rose.
I have also beaten them from oak, Hawthorn and Blackthorn.
Weigela. Upshire, North Essex
Guelder-rose. Fishers Green, Lee Valley, North Essex
Oak tree Potkiln Wood, Epping Forest, North Essex
During July 2020 Mark Hows and I were beating either side of a foot path at Fishers Green, Essex and we were finding Epaulet Ladybirds in almost all pieces of vegetation we sampled, including oak trees, thistles, burdock, White Bryony and Black Horehound.
Oak tree Fishers Green, Lee Valley, North Essex
White Bryony Fishers Green, Lee Valley, North Essex
